Page:History of botany (Sachs; Garnsey).djvu/282

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Examination of the Matured Framework
[Book II.

phenomena of vegetative life, that the true cell-body, the cell=protoplasm is prior in time and in conception, and can claim the higher position.




Mirbel, to whom we now return, had in 1801 laid down a theory of cell-formation which agreed in the main with that of Caspar Friedrich Wolff; he supposed with Wolff that each cell-space was separated from its neighbour by a single wall, and relying on fresh observations asserted the existence of visible pores in the dividing walls of parenchyma and of vessels, and also maintained some new views on the nature and formation of vessels. The essential points of this theory found an opponent in Germany in the person of Kurt Sprengel, the well-known historian of botany, and one of the most variously accomplished botanists of his time, who had published in 1802 a work written in diffuse and familiar style under the title of 'Anleitung zur Kenntniss der Gewächse.' He relied on his own observations, but these were evidently made with small magnifying powers, an obscure field of sight, and indifferent preparations. The cell-tissue, says Sprengel, consists of cavities of very various shape communicating with one another, the dividing walls being in some places broken through and in others wanting. He took the starch-granules which he saw in the seed-leaves of beans and other plants for vesicles, which increase in size by absorption of water and so form new tissue; but he did not explain how we are to conceive of the growth of organs with such a mode of cell-formation. His account of the vessels is extremely obscure, even more obscure than Hedwig's, though he has the merit of refuting the latter's strange theory of reconducting vessels in the epidermis; he also suggested, though only incidentally, the happy idea that spiral passages and even vessels might arise from cell-tissue, since the youngest parts of plants have only the latter; but he did not attempt to explain how or where the process takes place. Like Malpighi