Page:History of botany (Sachs; Garnsey).djvu/285

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Chap. III.]
of Cell-membrane in Plants.
265

mentions Hedwig's notion already described, and shows that its exact opposite is true, namely, that the spiral band is surrounded by a membrane on the outside, a fact which was afterwards denied by Link, Sprengel, and Moldenhawer. On the other hand he did not understand the sculpturing on the scalariform vessels; he took the pits in the dotted vessels for thickenings of the wall, such as are seen in the transverse ridges between the slits in true scalariform vessels, and the slits he thought were closed. If there was much that was erroneous in these views, yet Bernhardi contributed essentially to the clearing up of the subject by his effort to distinguish the different forms of air-vessels, and especially by pointing attention to the fact that neither spiral nor annular vessels are found in secondary wood. The resemblance between different forms of vessels misled many of his contemporaries into supposing that they are due to metamorphosis of true spiral vessels. Bernhardi showed that different forms of wall are found inside one vascular tube, but that this does not depend on modification with age; observation rather teaches that every kind of vessel receives its character in its young state, and especially that the youngest scalariform vessels do not present the form of spiral vessels.

Under the head of vessels proper he reckoned all tubular forms filled with a peculiar juice, milk-cells and true milk-vessels, and also resin-ducts and the like, and he made many good and still valuable observations on their distribution and sap-contents. He could not see the differences of structure in these various fluid-conveying vessels with the low magnifying power of his glass, and therefore attended chiefly to the structure of the large resin-ducts, which on the whole he rightly understood.

The question whether there are any other forms of vessels in the plant beside those already named gave him occasion to define a vessel better than it had yet been defined, namely as an uninterrupted tube or canal, and at the same time he found himself obliged to consider whether his bast-threads are vessels;