called the "nucleus;" while, at its hinder end, is a "contractile vacuole," conspicuous by its regular rhythmic appearances and disappearances. Obviously, although the Colpoda is not a monad, it differs from one only in subordinate details. Moreover, under certain conditions, it becomes quiescent, incloses itself in a delicate case or cyst, and then divides into two, four, or more portions, which are eventually set free and swim about as active Colpodæ.
But this creature is an unmistakable animal, and full-sized Colpodæ may be fed as easily as one feeds chickens. It is only needful to diffuse very finely ground carmine through the water in which they live, and, in a very short time, the bodies of the Colpodæ are stuffed with the deeply coloured granules of the pigment.
And if this were not sufficient evidence of the animality of Colpoda, there comes the fact that it is even more similar to another well-known ‘animalcule, Paramæcium, than it is to a monad. But Paramæcium is so huge a creature compared with those hitherto discussed — it reaches one one-hundred-and-twentieth of an inch or more in length — that there is no difficulty in making out its organization in detail; and in proving that it is not only an animal, but that it is an animal which possesses a somewhat complicated organization. For example, the surface layer of its body is different in structure from the deeper parts. There are two contractile vacuoles, from each of which radiates a system of vessel-like canals; and not only is there a conical depression continuous with a tube, which serve, as mouth and gullet, but the food ingested takes a definite course and refuse is rejected from a definite region. Nothing is easier than to feed these animals and to watch the particles of indigo or carmine accumulate at the lower end of the gullet. From this they gradually project, surrounded by a ball of water, which at length passes with a jerk, oddly simulating a gulp, into the pulpy central substance of the body, there to circulate up one side and down the other, until its contents are digested and assimilated. Nevertheless, this complex animal multiplies by division, as the monad does, and, like the monad, undergoes conjugation. It stands in the same relation to Heteromita on the animal side, as Coleochæte does on the plant side. Start from either, and such an insensible series of gradations leads to the monad that it is impossible to say at any stage of the progress — here the line between the animal and the plant must be drawn.
There is reason to think that certain organisms which pass through a monad stage of existence, such as the Myxomycetes, are, at one time of their lives, dependent upon external sources for their protein-matter, or are animals; and at another period manufacture it, or are plants. And seeing that the whole progress of modern investigation is in favour of the doctrine of continuity, it is a fair and probable speculation — though only a speculation — that, as there are some plants which can manufacture protein out of such apparently intractable mineral matters as carbonic acid, water, nitrate of ammonia, and metallic salts; while others need to be supplied with their carbon and nitrogen in the somewhat less raw form of tartrate of ammonia and allied compounds; so there may be yet others, as is possibly the case with the true parasitic plants, which can only manage to put together materials still better prepared — still more nearly approximated to protein — until we arrive at such organisms as the Psorospermiæ and the Panhistophyton, which are as much animal as vegetable in structure, but are animal in their dependence on other organisms for their food.
The singular circumstance observed by Meyer, that the Torula of yeast, though an indubitable plant, still flourishes most vigorously when supplied with the complex nitrogenous substance, pepsin; the probability that the Peronospora is nourished directly by the protoplasm of the potato-plant; and the wonderful facts which have recently been brought to light respecting insectivorous plants, all favour this view; and tend to the conclusion that the difference between animal and plant is one of degree rather than of kind; and that the problem whether, in a given case, an organism is an animal or a plant, may be essentially insoluble.
HER DEAREST FOE.
CHAPTER XXXIX.
Welcome as he ever was, Tom Reed was perhaps never so anxiously looked for as on the present occasion. Kate felt that he could disentangle the ravelled skein of her affairs; that he only could deal with Trapes; and his tact so manipulate the difficulties with which her relations
