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ON ACTIVE MOLECULES.
469

In Periploceæ, and in a few Apocineæ, the pollen, which in these plants is separable into componnd grains filled with spherical moving particles, is applied to processes of the stigma, analogous to those of Asclepiadeæ. A similar economy exists in Orchideæ, in which the pollen masses are always, at least in the early stage, granular; the grains, whether simple or compound, containing minute, nearly spherical particles, but the whole mass being, with [7 very few exceptions, connected by a determinate point of its surface with the stigma, or a glandular process of that organ.

Having found motion in the particles of the pollen of all the living plants which I had examined, I was led next to inquire whether this property continued after the death of the plant, and for what length of time it was retained.

In plants, either dried or immersed in spirit for a few days only, the particles of pollen of both kinds were found in motion equally evident with that observed in the living plant; specimens of several plants, some of which had been dried and preserved in an herbarium for upwards of twenty years, and others not less than a century, still exhibited the molecules or smaller spherical particles in considerable numbers, and in evident motion, along with a few of the larger particles, whose motions were much less manifest, and in some cases not observable.[1]

In this stage of the investigation having found, as I believed, a peculiar character in the motions of the particles of pollen in water, it occurred to me to appeal to this peculiarity as a test in certain families of Cryptogamous plants, namely, Mosses, and the genus Equisetum,

  1. While this sheet was passing through the press I have examined the pollen of several flowers which have been immersed in weak spirit about eleven months, particularly of Viola tricolor, Zizania aquatica, and Zea Mays; and in all these plants the peculiar particles of the pollen, which are oval or short oblong, though somewhat reduced in number, retain their form perfectly, and exhibit evident motion, though I think not so vivid as in those belonging to the living plant. In Viola tricolor, in which, as well as in other species of the same natural section of the genus, the pollen has a very remarkable form, the grain on immersion in nitric acid still discharged its contents by its four angles, though with less force than in the recent plant.