margin project in the same plane with the leaf, or more commonly (see Fig. 2) are considerably reflexed.
If a small object be placed on the glands in the centre of the leaf, a motor impulse is transmitted to the marginal tentacles. The nearer ones are first affected, and then those farther off, until at last all are slowly but unerringly inflected, and close over the object. This takes place in from one to five or more hours; the difference in time depending
Fig. 2.—Drosera rotundefolia.—Old leaf viewed laterally; enlarged about five times.
on several circumstances, as the size of the object and its nature; on the vigor and age of the leaf; whether it has lately been in action; and the temperature.
The tentacles in the centre do not become inflected when directly excited, though they are capable of inflection if excited by a motor impulse from other glands; but through and from them the motor impulse spreads gradually on all sides. Such is not the case with the marginal tentacles. If a bit of meat be placed on one of these it quickly transmits an impulse to its own bending portion, but never to those adjoining (see Fig. 5), for these are never affected until the meat has been carried to the central glands, which then radiate their conjoined impulse on all sides.
The sensitiveness of the leaves is located in the glands together with the immediately underlying cells of the tentacles. Though it is necessary that the glands should be touched, it is wonderful how slight a pressure will suffice. A bit of human hair 150 of an inch in length and weighing only 178740 of a grain will induce motion, transmit a motor impulse through the whole length of a marginal tentacle, and cause it to sweep through an angle of 180° or more. This minute morsel, it must be borne in mind, rests upon and is supported by the dense, viscid fluid which surrounds the gland, and the pressure is thus rendered inconceivably slight. Mr. Darwin conjectures that it may be less than the millionth of a grain. While the pressure may be extremely slight, it needs must be steady. A sharp, sudden brush of the tentacles does not induce inflection, nor do drops of water falling upon the glands from any height. This specialized nature of the sensitiveness may readily be seen to be of great use to the plant, effecting an economy of time and energy, for the process of inflection is slow and that of reëxpansion still slower, often occupy-
