Page:Wayside and Woodland Blossoms.djvu/267

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LICHENS.
128

walls, old posts and palings, on thatch and on the ground. Wherever they are found they may be accepted as certificates of the purity of the air. Formerly considered as a distinct type, they are now held by the advanced school of cryptogam ic botanists as commensals, or partnerships formed between a fungus and an alga. They are usually thin crusts, consisting of an upper and a lower epidermis, formed of closely crowded cells, and to the lower layer rootlike filaments are attached. Between these layers are two differing elements; a loose stratum of green cells (gonidia0 which are said to be algæ, and below these a layer of fungoid threads. The contention of the new school is that these algæ have been captured by a fungus and held in bondage, being forced to elaborate starch by means of their chlorophyll from the inorganic material obtained by the rootlike filaments, which starch the fungus is able to feed upon. Some of the green cells are pushed out from time to time invested with a few wisps of fungus-threads, and so reproduce the partnership. It is but right to add that some good authorities on this branch of botany decline to accept these views, and still regard lichens as independent organisms and not partnerships.

The species are very numerous, but their identification is not easy, and requires serious application. The two figured are exceedingly common in some districts. Various species of Cup-moss (Cladonia) will be met on heaths, sandy hedge-banks, etc. They have a flat crust-like base, from which arise pale grey tubes or cups, bearing at their tips the bright scarlet, pinky-brown, or even black fruits. A more common form in woods and on banks is Cladonia pyxidata, with the tube greatly increasing in width upwards. Cladonia rangiferina is the well-known Reindeer-moss, of inestimable value in extreme Northern latitudes as the food of the useful animal whose name it bears; it may be found in abundance in this country on heaths and hillsides covering the ground beneath the heather.