The side of the skull, in front of the operculum, is covered by a large irregularly-shaped bone (T) (corresponding to the "tympanic lamina" of Ceratodus, Fig. 35, q), held by some to be the præoperculum; along its upper circumference lies a series of small ossicles, of which two may be distinguished as spiraculars, as they form a valve for the protection of the spiracular orifice of these fishes. An infraorbital ring is represented by a præ- and post-orbital only.
Each hyoid consists of three pieces, none of which bear branchiostegals, the single median piece being osseous in front and cartilaginous behind. Four branchial arches are developed, the foremost consisting of three, the second and third of two, and the last of a single piece. There is no lower pharyngeal. Between the rami of the lower jaw the throat is protected by a pair of large osseous laminæ (gular plates), which have been considered to represent the urohyal of osseous fishes.
The scapulary arch is almost entirely formed by the well-developed membrane-bones, which in the ventral line are suturally united. The pectoral fin is supported by three bones, pro-, meso-, and metapterygium, of which the dilated middle one alone bears rays, and is excluded from the articulation with the shoulder-girdle.
The pubic consists of paired bone, to which tarsal bones supporting the fin-rays are attached.
In the Lepidosteoidei the vertebræ are completely ossified, and opisthocœlous, having a convexity in front and a concavity behind, as in some Amphibians. Though the end of the body externally appears nearly diphycercal, the termination of the vertebral column is, in fact, distinctly heterocercal (Fig. 40). Its extremity remains cartilaginous, is turned upwards, and lies immediately below the scutes which cover the upper margin of the caudal fin. It is preceded by a few rudimentary vertebræ which gradually pass into the fully developed normal vertebræ. The caudal fin is suspended from hæmapophyses
