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The Descent of Man.
Part I.

confirmed by Prof. Kupffer. M. Kovalevsky writes to me from Naples, that he has now carried these observations yet further; and should his results be well established, the whole will form a discovery of the very greatest value. Thus, if we may rely on embryology, ever safest guide in classification, it seems that we have at last gained a clue to the source whence the Vertebrata were derived.[1] We should then be justified in believing that at an extremely remote period a group of animals existed, resembling in many respects the larvæ of our present Ascidians, which diverged into two great branches—the one retrograding in development and producing the present class of Ascidians, the other rising to the crown and summit of the animal kingdom by giving birth to the Vertebrata.

We have thus far endeavoured rudely to trace the genealogy of the Vertebrata by the aid of their mutual affinities. We will now look to man as he exists; and we shall, I think, be able partially to restore the structure of our early progenitors, during successive periods, but not in due order of time. This can be effected by means of the rudiments which man still retains, by the characters which occasionally make their appearance in him through reversion, and by the aid of the principles of morphology and embryology. The various facts, to which I shall here allude, have been given in the previous chapters.

The early progenitors of man must have been once covered with hair, both sexes having beards; their ears were probably pointed, and capable of movement; and their bodies were provided with a tail, having the proper muscles. Their limbs and bodies were also acted on by many muscles which now only occasionally reappear, but are normally present in the Quadrumana. At this or some earlier period, the great artery and nerve of the humerus ran through a supra-condyloid foramen. The intestine gave forth a much larger diverticulum or cæcum than that now existing. The foot was then prehensile, judging from the condition of the great toe in the fœtus; and our progenitors, no doubt, were arboreal in their habits, and frequented some warm, forest-clad land. The males had great canine teeth, which

  1. But I am bound to add that some competent judges dispute this conclusion; for instance, M. Giard, in a series of papers in the 'Archives de Zoologie Expérimentale,' for 1872. Nevertheless, this naturalist remarks, p. 281, "L’organisation de la larve ascidienne en dehors de toute hypothèse et de toute théorie, nous montre comment la nature peut produire la disposition fondamentale du type vertébré (l’existence d’une corde dorsale) chez un invertébré par la seule condition vitale de l’adaptation, et cette simple possibilité du passage supprime l’abîme entre les deux sous-règnes, encore bien qu’en ignore par où le passage s’est fait en réalité."