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354
The Descent of Man.
Part II.

attack them. Nor can I quite disbelieve the several accounts which have appeared of their thus paralysing their prey with fear. Some other snakes also make a distinct noise by rapidly vibrating their tails against the surrounding stalks of plants; and I have myself heard this in the case of a Trigonocephalus in S. America.

Lacertilia.—The males of some, probably of many kinds of lizards fight together from rivalry. Thus the arboreal Anolis cristatellus of S. America is extremely pugnacious: "During the spring and early part of the summer, two adult males rarely meet without a contest. On first seeing one another, they nod their heads up and down three or four times, and at the same time expanding the frill or pouch beneath the throat; their eyes glisten with rage, and after waving their tails from side to side for a few seconds, as if to gather energy, they dart at each other furiously, rolling over and over, and holding firmly with their teeth. The conflict generally ends in one of the combatants losing his tail, which is often devoured by the victor." The male of this species is considerably larger than the female;[1] and this, as far as Dr. Günther has been able to ascertain, is the general rule with lizards of all kinds. The males alone of the Cyrtodactylus rubidus of the Andaman Islands possesses pre-anal pores; and these pores judging from analogy probably serve to emit an odour.[2]

The sexes often differ greatly in various external characters. The male of the above-mentioned Anolis is furnished with a crest which runs along the back and tail, and can be erected at pleasure; but of this crest the female does not exhibit a trace. In the Indian Cophotis ceylanica, the female has a dorsal crest, though much less developed than in the male; and so it is, as Dr. Günther informs me, with the females of many Iguanas, Chameleons, and other lizards. In some species, however, the crest is equally developed in both sexes, as in the Iguana tuberculata. In the genus Sitana, the males alone are furnished with a large throat-pouch (fig. 33), which can be folded up like a fan, and is coloured blue, black, and red; but these splendid colours are exhibited only during the pairing-season. The female does not possess even a rudiment of this appendage. In the Anolis cristatellus, according to Mr. Austen, the throat pouch, which is bright red marbled with yellow, is present in the female, though in a rudimental condition. Again, in certain other lizards, both sexes are equally well provided with throat pouches. Here we

  1. Mr. N. L. Austen kept these animals alive for a considerable time; see 'Land and Water,' July 1867, p. 9.
  2. Stoliczka, 'Journal of Asiatic Soc. of Bengal,' vol. xxxiv. 1870, p. 166.