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The Descent of Man.
Part II.

racter. Thus in the male of our common peewit (Vanellus cristatus) the tubercle on the shoulder of the wing becomes more prominent during the breeding-season, and the males fight together. In some species of Lobivanellus a similar tubercle becomes developed during the breeding-season "into a short horny spur." In the Australian L. lobatus both sexes have spurs, but these are much larger in the males than in the females. In an allied bird, the Hoplopterus armatus, the spurs do not increase in size during the breeding-season; but these birds have been seen in Egypt to fight together, in the same manner as our peewits, by turning suddenly in the air and striking sideways at each other, sometimes with fatal results. Thus also they drive away other enemies.[1]

The season of love is that of battle; but the males of some birds, as of the game-fowl and ruff, and even the young males of the wild turkey and grouse,[2] are ready to fight whenever they meet. The presence of the female is the teterrima belli causa. The Bengali baboos make the pretty little males of the amadavat (Estrelda amandava) fight together by placing three small cages in a row, with a female in the middle; after a little time the two males are turned loose, and immediately a desperate battle ensues.[3] When many males congregate at the same appointed spot and fight together, as in the case of grouse and various other birds, they are generally attended by the females,[4] which afterwards pair with the victorious combatants. But in some cases the pairing precedes instead of succeeding the combat: thus according to Audubon,[5] several males of the Virginian goat-sucker (Caprimulgus virginianus) "court, in a highly entertaining manner the female, and no sooner has she made her choice, than her approved gives chase to all intruders, and drives


    Pala, Brehm's ' Thierleben,' B. iv. s. 740. See also on this bird Azara, 'Voyages dans l'Amérique mérid.' tom. iv. 1809, pp. 179, 253.

  1. See, on our peewit, Mr. R. Carr in ' Land and Water,' Aug. 8th, p. 46. In regard to Lobivanellus, see Jerdon's ' Birds of India,' vol. iii. p. 647, and Gould's 'Handbook of Birds of Australia,' vol. ii. p. 220. For the Holopterus, see Mr. Allen in the ' Ibis,' vol. v. 1836 , p. 156.
  2. Audubon, 'Ornith. Biography,' vol. ii. p. 492; vol. i. pp. 4–13.
  3. Mr. Blyth, 'Land and Water, 1867 , p. 212.
  4. Richardson on Tetrao umhellus 'Fauna Bor, Amer.: Birds,' 1831, p. 343. L. Lloyd, 'Game Birds of Sweden,' 1867, pp. 22, 79, on the capercailzie and black-cock. Brehm, however, asserts ('Thierleben,' &c., B. iv. s. 352) that in Germany the grey-hens do not generally attend the Balzen of the black-cocks, but this is an exception to the common rule; possibly the hens may lie hidden in the surrounding bushes, as is known to be the case with the grey-hens in Scandinavia, and with other species in N. America.
  5. 'Ornithological Biography,' vol. ii. p. 275.