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COLOMBIA
[FAUNA AND FLORA


the equator into the south torrid zone. The great Andean barrier which crosses the republic from the south to north acts as a condenser to the prevailing easterly winds from the Atlantic, and causes a very heavy rainfall on their eastern slopes and over the forested Amazon plain. High temperatures as well as excessive humidity prevail throughout this region. Farther north, on the open llanos of the Orinoco tributaries, the year is divided into equal parts, an alternating wet and dry season, the sun temperatures being high followed by cool nights, and the temperatures of the rainy season being even higher. The rainfall is heavy in the wet season, causing many of the rivers to spread over extensive areas, but in the dry season the inundated plains become dry, the large rivers fed by the snows and rainfall of the Andes return within their banks, the shallow lagoons and smaller streams dry up, vegetation disappears, and the level plain becomes a desert. The northern plains of the republic are swept by the north-east trades, and here, too, the mountain barriers exercise a strongly modifying influence. The low ridges of the Sierra de Perijá do not wholly shut out these moisture-laden winds, but they cause a heavy rainfall on their eastern slopes, and create a dry area on their western flanks, of which the Vale of Upar is an example. The higher masses of the Sierra Nevada de Santa Marta cover a very limited area, leaving the trade winds a comparatively unbroken sweep across the northern plains until checked by the Western Cordillera, the Panama ranges and the Sierra de Baudo, where a heavy precipitation follows. Farther south the coast ranges cause a very heavy rainfall on their western slopes, which are quite as uninhabitable because of rain and heat as are the coasts of southern Chile through rain and cold. The rainfall on this coast is said to average 73 in., though it is much higher at certain points and in the Atrato Valley. As a result the coastal plain is covered with swamps and tangled forests, and is extremely unhealthy, except at a few favoured points on the coast. High temperatures prevail throughout the greater part of the Magdalena and Cauca valleys, because the mountain ranges which enclose them shut out the prevailing winds. At Honda, on the Magdalena, 664 ft. above sea-level, the mean temperature for the year is 82° F., and the mercury frequently rises to 102° in the shade. These lowland plains and valleys comprise the climatic tropical zone of Colombia, which is characterized by high temperatures, and by excessive humidity and dense forests, an exception to the last-named characteristic being the open llanos where dry summers prevail. Above this tropical zone in the mountainous regions are to be found all the varying gradations of climate which we are accustomed to associate with changes in latitude. There are the subtropical districts of the valleys and slopes between 1500 and 7500 ft. elevation, which include some of the most fertile and productive areas in Colombia; the temperate districts between 7500 and 10,000 ft., the cold, bleak and inhospitable paramos between 10,000 and 15,000 ft., and above these the arctic wastes of ice and snow. The temperate and subtropical regions cover the greater part of the departments traversed by the Eastern Cordillera, the northern end of the Central Cordillera, the Santa Marta plateaus, and the Upper Cauca Valley. They include the larger part of the white population and the chief productive industries of the country. There is no satisfactory record of temperatures and rainfall in these widely different climatic zones from which correct averages can be drawn and compared. Observations have been made and recorded at Bogotá and at some other large towns, but for the greater part of the country we have only fragmentary reports. The mean annual temperature on the eastern plains, so far as known, ranges from 87° F. on the forested slopes to 90° and 91° on the llanos of the Meta and Arauca. On the Caribbean coastal plain it ranges from 80° to 84°, but at Tumaco, on the Pacific coast, within two degrees of the equator, it is only 79°. At Medellin, in the mountainous region of Antioquia, 4950 ft. above sea-level, the mean annual temperature is 70°, and the yearly rainfall 55 in., while at Bogotá, 8563 ft., the former is 57° and the latter 44 in. At Tuquerres, near the frontier of Ecuador, 10,200 ft. elevation, the mean annual temperature is said to be 55°. The changes of seasons are no less complicated and confusing. A considerable part of the republic is covered by the equatorial belt of calms, whose oscillations divide the year into a wet and dry season. This division is modified, however, by the location of mountain ranges and by elevation. In the Amazon region there is no great change during the year, and on the northern plains the so-called dry season is one of light rains except where mountain ranges break the sweep of the north-east trades. The alternating wet and dry seasons are likewise to be found on the Pacific coastal plain, though this region is not entirely dry and vegetation never dries up as on the llanos. Above the lowland plains the seasons vary in character according to geographical position and elevation. The two-season division rules in the departments of Santander and Antioquia, but without the extremes of humidity and aridity characteristic of the eastern plains. Farther south, at elevations between 800 and 9500 ft., the year is divided into four distinct seasons—two wet and two dry—the former called inviernos (winters) and the latter veranos (summers). These seasons are governed by the apparent movements of the sun, the winters occurring at the equinoxes and the summers at the solstices. The sabana of Bogotá and neighbouring districts are subject to these changes of season. At higher altitudes long, cold, wet winters are experienced, with so short and cold a summer between them that the bleak paramos are left uninhabited except by a few shepherds in the short dry season.

Fauna.—The geographical position of Colombia gives to it a fauna and flora largely characteristic of the great tropical region of the Amazon on the south-east, and of the mountainous regions of Central America on the north-west. At the same time it is rich in animal and plant types of its own, especially the latter, and is considered one of the best fields in South America for the student and collector. The fauna is essentially tropical, though a few species characteristic of colder regions are to be found in the higher Andes. Of the Quadrumana there are at least seventeen distinct species, and this number may be increased after a thorough exploration of the forested eastern plains. They are all arboreal in habit, and are to be found throughout the forested lowlands and lower mountain slopes. The carnivora are represented by seven or eight species of the Felidae, the largest of which are the puma (Felis concolor) and the jaguar (F. onca). These animals, together with the smaller ocelot, have a wide geographical range, and are very numerous in the valley of the Magdalena. Two species of bear and the “coatí” (Nasua) represent the plantigrades and inhabit the mountain slopes, and, of Pachydermata, the peccary (Dicotyles) and “danta” or tapir (Tapirus) have a wide distribution throughout the lowland and lower plateau forests. The Colombian tapir is known as the Tapirus Roulini, and is slightly smaller than the Brazilian species (T. americanus). There are deer in the forests and on the open savannahs, the rabbit and squirrel are to be seen on the eastern slopes of the Andes, and partly amphibious rodents, the “capybara” (Hydrochoerus) and “guagua” (Coelogenys subniger), are very numerous along the wooded watercourses. The sloth, armadillo, opossum, skunk and a species of fox complete the list of the more common quadrupeds so far as known, though it is certain that a careful biological survey would discover many others. The large rivers of Colombia and the lakes of the lowlands are filled with alligators, turtles, and fish, and several species of fish are highly esteemed by the natives as food. The saurians are represented on land by several species of lizard, some of them conspicuous for their brilliant colouring, and by the large “iguana,” whose flesh is considered a great delicacy. Among the ophidians, which include many harmless species, are the boa-constrictor, rattlesnake, the dreaded Lachesis and the coral snake. The “manatee” (Manatus americanus) is found in the Atrato and other large Colombian rivers.

In bird and insect life Colombia is second only to Brazil. The condor, which inhabits the higher Cordilleras, is peculiar to the whole Andean region, and is the largest of the Raptores. Among other members of this order are the eagle, osprey, vulture, buzzard, kite and hawk, with about a dozen species in all. Parrots and paroquets are numerous everywhere in the tropical and subtropical regions, as also the gorgeously coloured macaw and awkward toucan. The largest class, perhaps, is that formed by the astonishing number of water-fowl which throng the shallow lagoons and river beaches at certain seasons of the year. They are mostly migratory in habit, and are to be found in many other countries. Among these are the large white crane and small crane, the blue heron, the snowy-white egret, the roseate spoonbill (Platalea ajaja), stork, bittern and many species of ducks. The largest and most conspicuous member of this interesting family is the Mycteria americana, the gigantic stork so frequently seen in the Amazon valley, and even more numerous about the lagoons of northern Colombia. One of the best game-birds of the forest is the “crested curassow” (Crax alector), sometimes weighing 12lb, which feeds on arboreal fruits and rarely comes to the ground. Colombia also possesses many species of the beautiful little humming-bird, among which are the tiny Steganura Underwoodi and the sword-bill, Docimastes ensiferus, which were found by Mr Albert Millican on a bleak paramo 12,000 ft. above sea-level. One of the most interesting birds found in the country is the “weaver-bird” (Cassicus persicus), which lives in colonies and suspends its long, pouch-like nest from the end of a horizontal branch of some high, isolated tree. In regard to insects, what has been said of Brazil will apply very closely to Colombia. Mosquitoes, butterflies, spiders, beetles and ants are infinitely numerous, and some of the species are indescribably troublesome.

Flora.—The Colombian flora is richer in species and individual characteristics than the fauna, owing in part to its greater dependence on climatic conditions. It ranges from the purely tropical types of the lowlands to the Alpine species of the more elevated paramos. It should be remembered, however, that large areas of the lowland plains have only a very limited arboreal growth. These plains include the extensive llanos of the Orinoco tributaries where coarse, hardy grasses and occasional clumps of palms are almost the only vegetation to be seen. There are other open plains in northern Colombia, sometimes covered with a shrubby growth, and the “mesas” (flat-topped mountains) and plateaus of the Cordilleras are frequently bare of trees. Farther up, on the cold, bleak paramos, only stunted and hardy trees are to be found. On the other hand, a luxuriant forest growth covers a very large part of the republic, including the southern plains of the Amazon tributaries, the foothills, slopes and valleys of the Cordilleras, a larger part of the northern plains, and the whole surface of the Western Cordillera and coast. The most conspicuous and perhaps the most universal type in all these regions, below an approximate elevation of 10,000 ft.,