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probably resembled the bears more than any living animals, with the important exception that in their feet they were much like the elephant. To the general proportions of the bears must be added the tail of medium length. Whether they were covered with hair or not is of course uncertain. Of their nearest living allies, the elephants, some were hairy and others naked.... The movements of the Coryphodons doubtless resembled those of the elephant in its shuffling and ambling gait, and may have been even more awkward from the inflexibility of the ankle."

The most recent members of this sub-order come from the Middle Eocene beds, and are chiefly referable to the genus Dinoceras, with which Tinoceras and Uintatherium are at least very nearly related, if not identical. These creatures were of great size, larger than the earlier types which have been considered. They show a certain superficial resemblance to the Titanotheriidae, on account of the massive horn cores upon the skull. These horn cores are large upon the maxillae and the parietals, and are paired; on the nasals are smaller horns. The bones of the skull have air cavities. The incisors of the upper jaw are absent; the canines are enormous tusks, and the lower jaws are flanged downwards near the symphysis where these tusks border them. Contrary to what is found in the older types, where the position of the condyle of the lower jaw is normal, this prominence faces backwards in the Dinocerata. The same shortness of the spines of the dorsal vertebrae prevails in this group as in the other Amblypoda, though it is perhaps hardly so marked. The scapula has not the peculiar acuminate form that exists in Coryphodon, but is triangular and broad above. The limbs are elephantine, in that the angle between the humerus and the femur respectively, and the bones which follow, is not marked. The hind-limbs are especially straight. The tail is short as compared with that of the primitive Amblypoda. The Dinocerata are purely digitigrade. The entepicondylar foramen has, as in the Coryphodonts, disappeared. The os centrale of the carpus has become fused, and no longer exists as a separate bone. The fibula no longer articulates with the calcaneum, but both that bone and the ulna are well developed. The genus Astrapotherium is placed among the Amblypoda by some authorities.[1]

  1. Gadow, A Classification of Vertebrata, Recent and Extinct, London, 1898.