size than Anchitherium, to be considered immediately. The odontoid process of the axis is just beginning to assume the characteristic spout-like shape of that of the existing Horse and many modern Ungulates. The median digit of both fore- and hind-limbs has become greatly enlarged as compared with the corresponding digit of earlier forms.
It is held, however, that Anchitherium is not on the direct line of descent either in America or in Europe, in both of which it occurs. Its teeth are in some respects less Horse-like than in some of the more ancient genera, to which the converse would be expected on the descent theory. Its hoofs are much elongated and flattened, a mark of specialisation and not appropriate to a creature holding an intermediate position in the equine series. Both the American (A. equinum) and the European species (A. aureliense) are of very large size, larger than its successors, and such "alternations in bulk are unlikely."
The genus Desmatippus of Professor Scott[1] fills in the gap between Miohippus and Protohippus. The molars and premolars are brachyodont, but there is a thin deposit of cement in the tooth valleys, leading towards the more complete filling of these valleys with cement, which is found in Protohippus. This genus of Horses, of which there is at present but one species, D. crenidens, was three-toed, and "the lateral digits, so far as can be judged by fragmentary remains, were still fairly developed, and though much more reduced than in Miohippus, appear to be somewhat less so than in Protohippus."
To recapitulate, the following is the probable series of equines in America—Mesohippus, Miohippus, Desmatippus, Protohippus.
The development of the limbs of the Horse shows a most interesting series, of stages, which correspond in part to the ancestral forms which palaeontology seems to prove to be the line of the descent of our existing Equidae. This matter has recently been elucidated by Professor Ewart, who details the following facts and comparisons:—
In the youngest embryo (about 20 mm. in length) the humerus is somewhat curved, and considerably longer than the radius and carpus taken together. The first-named bone is shorter in the adult, and the proportions of that bone in the young as well as its curvature are suggestive of that ancient
- ↑ Trans. American Phil. Soc. xviii. 1896, p. 55.
