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instead of rudimentary like those of other Whales, the blow-hole lies more in the middle of the face. The skull, too, is not Whale-like in a number of other points. Thus the premaxillaries take their fair share in the outline of the upper jaw; and, furthermore, bear the incisor teeth. The parietals meet above in a crest and are not excluded from the roof of the skull. The vertebrae of the neck are in no way shortened; neither are they fused together. The ribs are double-headed, and the sternum is made up of several pieces. Some naturalists, particularly Professor D'Arcy Thompson,[1] have assigned a relationship to the Seals to these ancient Cetacea; but others[2] have disputed this view chiefly on the grounds that the characters which appear to be Seal-like are simply characters which are generalised and so far at most not Whale-like. Thus the long neck and the serrated character of the teeth may be accepted as Seal-like on the one hand; but on the other, a simple serrated tooth and a long neck are not by any means features of organisation which we should consider out of the way in an ancient form of Cetacean which probably preyed upon fish. The humerus of Zeuglodon, according to Mr. Lydekker, puts out of court any possible near relationship to the Seals. But the matter under dispute can be further studied by reference to the three memoirs quoted below.

  1. Thompson, Studies Mus. Dundee, i. 1890; and C. R. Congrès de Zoologie, 1889, p. 225.
  2. Lydekker, Proc. Zool. Soc. 1892, p. 560.