Page:Manual of the New Zealand Flora.djvu/628

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POLYGONACEÆ.
[Polygonom.

spikes or racemes, often forming terminal panicles. Perianth inferior, persistent; segments 3–6, free or connate at the base, imbricate. Stamens 5–9, rarely more or less, hypogynous or perigynous; filaments capillary or subulate, free or connate at the base; anthers 2-celled. Ovary superior, compressed or 3-gonous; styles 1–3; ovule solitary, basal, orthotropous. Fruit a small hard indehiscent trigonous or compressed nut, usually enclosed in the persistent perianth. Seed erect, testa membranous; albumen copious, farinaceous; embryo variable in position, radicle superior.

A rather large order, spread over the whole world, but most abundant in the north temperate zone. Genera 30; species over 600. The roots of many species are nauseous and purgative, the medicinal rhubarb being a well-known instance. On the other hand, the stems and leaves are frequently acid and refreshing, as the garden-rhubarb and the common sorrel. Buckwheat (Fagopyrum) is the only edible plant of any consequence. Of the 3 New Zealand genera, Muehlenbeckia extends to Australia and South America; the remaining 2 are widely distributed in both hemispheres.

Flowers hermaphrodite. Perianth-segments 5, not succulent in fruit 1. Polygonum.
Flowers hermaphrodite. Perianth-segments 6, the 3 inner enlarging and closing over the fruit, not succulent 2. Rumex.
Flowers unisexual. Perianth-segments 5, enlarged and succulent in fruit 3. Muehlenbeckia.


1. POLYGONUM, Linn.

Herbs, rarely shrubby at the base. Leaves alternate; stipules membranous, usually tubular and closely sheathing the stem. Flowers small, hermaphrodite, clustered; clusters either axillary or in racemes or spikes or panicles; bracts and bracteoles membranous, ochreate; pedicels usually jointed. Perianth 5-partite, green or coloured; segments equal or the 2 or 3 outer ones the largest. Stamens 5–8. Ovary compressed or 3-gonous; styles 2 or 3, free or connate at the base; stigmas usually capitellate. Nuts compressed or 3-gonous, included in the persistent perianth. Seed albuminous; embryo excentric or lateral; radicle long, superior.

A large and almost cosmopolitan genus, including over 150 species. The three found in New Zealand are all widely distributed.

* Stems prostrate. Flowers in axillary clusters.
Stems long, wiry, sparingly branched. Perianth ⅛ in. long. Nut minutely striate or punctate 1. P. aviculare.
Stems short, compactly branched. Perianth 1/12 in. long. Nut smooth and polished 2. P. plebeium.
colspan=2 ** Stems erect or ascending above. Flowers in slender terminal spikes.
Leaves 2–5 in., lanceolate. Stipules ciliate and pilose. Spikes very slender, 1–2 in. long 3. P. serrulatum.

For references to several introduced species of Polygonum, see the list of naturalised plants appended to this work.