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14
The Problems

remembered that the principle is only explicitly declared to apply to discontinuous characters[1]. As stated also it can only be true where reciprocal crossings lead to the same result. Moreover, it can only be tested when there is no sensible diminution in fertility on crossing.

Upon the appearance of de Vries' paper announcing the "rediscovery" and confirmation of Mendel's law and its extension to a great number of cases two other observers came forward almost simultaneously and independently described series of experiments fully confirming Mendel's work. Of these papers the first is that of Correns, who repeated Mendel's original experiment with Peas having seeds of different colours. The second is a long and very valuable memoir of Tschermak, which gives an account of elaborate researches into the results of crossing a number of varieties of Pisum sativum. These experiments were in many cases carried out on a large scale, and prove the main fact enuntiated by Mendel beyond any possibility of contradiction. The more exhaustive of these researches are those of Tschermak on Peas and Correns on several varieties of Maize. Both these elaborate investigations have abundantly proved the general applicability of Mendel's law to the character of the plants studied, though both indicate some few exceptions. The details of de Vries' experiments are promised in the second volume of his most valuable Mutationstheorie. Correns in regard to Maize and Tschermak in the case of P. sativum have obtained further proof that Mendel's law holds as well in the case of varieties differing from each other in two pairs of characters, one of each pair being dominant, though of course a more complicated expression is needed in such cases.[2]

  1. See later.
  2. Tschermak's investigations were besides directed to a re-exami-