Page:Mendel's principles of heredity; a defence.pdf/52

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32
The Problems

If however instead of pure extreme varieties we were to take a pair of varieties differing normally by only a foot or two, we might, owing to the masking effects of conditions, &c., have great difficulty in distinguishing the three forms. in the second generation. There would besides be twice as many heterozygous individuals as homozygous individuals of each kind, giving a symmetrical distribution of heights, and who might not—in pre-Mendelian days—have accepted such evidence—made still less clear by influence of conditions—as proof of Continuous Variation both of zygotes and gametes?

Suppose, then, that instead of two pure types, we had six or eight breeding together, each pair forming their own heterozygote, there would be a very remote chance of such purity or fixity of type whether of gamete or zygote being detected.

Dominance, as we have seen, is merely a phenomenon incidental to specific cases, between which no other common property has yet been perceived. In the phenomena of blended inheritance we clearly have no dominance. In the cases of alternative inheritance studied by Galton and Pearson there is evidently no universal dominance. From the tables of Basset hound pedigrees there is clearly no definite dominance of either of the coat-colours. In the case of eye-colour the published tables do not, so far as I have discovered, furnish the material for a decision, though it is scarcely possible the phenomenon, even if only occasional, could have been overlooked. We must take it, then, there is no sensible dominance in these cases; but whether there is or is not sensible gametic purity is an altogether different question, which, so far as I can judge, is as yet untouched. It may perfectly well be that we shall be compelled to recognize that in many cases there is no such purity, and