in many Compositæ, especially Cinarocephalæ. On the contrary, in Goodenoviæ the stigma at the same period is hardly visible, and is certainly not then capable of receiving impregnation from the pollen of its proper flower; it is therefore either impregnated by the antheræ of different flowers, or in some cases at a more advanced stage by the pollen of its own antheræ, which is received and detained in the indusium. To these arguments for the exclusion of Lobelia I may add that in the greater part of Goodenoviæ with dehiscent fruit the dissepiment is parallel to the valves of the capsule, in which respect they differ equally from Lobelia and the valvular-fruited Campanulaceæ; and lastly, that many species of Lobelia as well as Campanulaceae contain a milky juice of which there is no instance in Goodenoviæ. If, therefore, in Lobelia the pencil surrounding the stigma and the irregularity of the corolla, which, however, in some species is hardly perceptible, be considered as characters sufficient to separate this extensive genus from Campanulaceæ, it may form a separate order, admitting, perhaps, of subdivision into several distinct genera.
I have formerly observed[1] that in two genera of Goodenoviæ, namely, Euthales and Velleia, the base of the corolla coheres with the ovarium while the calyx remains entirely distinct. This structure I had stated as being peculiar to these genera, and as in some degree invalidating one of Jussieu's arguments for considering the floral envelope of Monocotyledones as calyx rather than corolla. The fact, however, seems not to be admitted by Richard, who in the dissertation already quoted[2] describes what has hitherto been called calyx in Velleia as bracteæ; a view of the structure which in those species of that genus having triphyllous calyx may appear plausible, but of which the probability is [561 diminished even in those with pentaphyllous calyx, and still more in Euthales, where the calyx is also tubular. But a stronger argument for the part usually denominated calyx being in these genera really such may be derived from certain species of Goodenia, in which it will be admitted that both calyx and corolla are present, and where, though
- ↑ Prodr. fl. Nov. Holl. 580.
- ↑ Annales du. mus. 18, p. 27.
