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ON THE ORGANS AND MODE OF FECUNDATION

Thirdly, I have never been able to find those perforations, represented by Mr. Bauer, in the bursiculæ of Orchis and Ophrys, and the existence of which in these genera is essential to his hypothesis.

And, lastly, the appearance of the stigma in Bletia Tan- 695] kervilliæ, after impregnation, as he believes, according to my view of the subject would rather prove that it was in a state capable of acting upon, but had not yet received the fecundating matter from, the anthera.

In thus venturing to differ from so accurate and experienced an observer as Mr. Bauer on a subject which he has for many years minutely studied and so beautifully illustrated, I am well aware how great a risk I incur of being myself found in error.

I am very desirous, however, that the perusal of this sketch of the various statements that have appeared on the question of impregnation, with the greater part of which he is at present probably unacquainted, should induce him to re-examine the facts and arguments by which his own opinion on this subject is supported. He will thus either succeed in establishing his theory on more satisfactory grounds, or, if the examination should prove unfavourable, he will, I am persuaded, from his well-known candour, as readily abandon it.

The notice here given of the opinions of botanists on impregnation in Orchideæ brings the subject down to the spring of the present year, when from circumstances, which I may hereafter have occasion to advert to, my attention was directed to this family of plants, the particular study of which I had for a long time discontinued.

In reviewing notes respecting them, made many years ago, I found some points merely hinted at, or imperfectly made out, which seemed deserving of further examination; and in the course of these inquiries, other observations of at least equal importance suggested themselves.