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14
The Romanes Lecture 1905

surface, is practically full, that is to say, fully occupied. Only one pair of young can grow up to take the place of the pair—male and female—which have launched a dozen, or it may be as many as a hundred thousand, young individuals on the world. The property of Variation ensures that amongst this excess of young there are many differences. Eventually those survive which are most fitted to the special conditions under which this particular organism has to live. The conditions may, and indeed in long lapses of time must, change, and thus some variation not previously favoured will gain the day and survive. The 'struggle for existence' of Darwin is the struggle amongst all the superabundant young of a given species, in a given area, to gain the necessary food, to escape voracious enemies, and gain protection from excesses of heat, cold, moisture, and dryness. One pair in the new generation—only one pair—survive for every parental pair. Animal population does not increase: 'Increase and multiply' has never been said by Nature to her lower creatures. Locally, and from time to time, owing to exceptional changes, a species may multiply here and decrease there; but it is important to realize that the 'struggle for existence' in Nature—that is to say, among the animals and plants of this earth untouched by man—is a desperate one, however tranquil and peaceful the battle-field may appear to us. The struggle for existence takes place, not as a clever French writer3 glibly informs his readers, between different species, but between individuals of the same species, brothers and sisters and cousins. The struggle between a beast of prey which seeks to nourish itself and the buffalo which defends its life with its horns is not 'the struggle for existence' so named by Darwin. More-