matin. Professor Wilson has brought forward the convincing data showing that the complex character of sex in insects actually resides in or is determined by particular and definite masses of this wonderful basis of inheritance.
Mendel's principles also account in the most remarkable way for many previously obscure phenomena, such as reversion, and again, the case where a child resembles its grandparent more than either of its parents; these seem to be due, so to speak, to the rise to the surface of a hidden stream of germ-plasm that had flowed for one or many generations beneath its accompanying currents. I believe that the law is replacing more and more the laws of Galton and Pearson, formulated as statistical summaries of certain phenomena of human inheritance taken en masse. According to Galton's celebrated law of ancestral inheritance, the qualities of any organism are determined to the extent of a certain fraction by its two parents taken together as a mid-parent, that a smaller definite fraction is contributed by the grandparents taken together as a mid-grandparent, and so on to earlier generations. But Mendel's Law has far greater definiteness, it explains more accurately the cases of alternative inheritance, and it may be shown to hold for blended and mosaic inheritance as well.
De Vries's mutation theory has already been explained in an earlier address by Professor Richards. It is clearly not an alternative but a complementary theory to natural selection, the germ-plasm and Mendelian theories. Like these last, it emphasizes the importance of the congenital hereditary qualities contained in the germ-plasm, though unlike the Darwinian doctrine it shows that sometimes new forms may arise by sudden leaps and not necessarily by the slow and gradual accumulation of slight modifications or fluctuations. The mutants like any other variants must present themselves before the jury of environmental circumstances, which passes judgment upon their condition of adaptation, and they, too, must abide by the verdict that means life or death.
From what has been said of these post-Darwinian discoveries, the Lamarckian doctrine, which teaches that acquired non-congenital characters are transmitted, seems to be ruled out. I would not lead you to believe that the matter is settled. I would say only that the non-transmission of racial mutilations, negative breeding experiments upon mutilated rats and mice, the results of further study of supposedly transmitted immunity to poisons—that all these have led zoologists to render the verdict of "not proved." The future may bring to light positive evidence, and cases like Brown-Séquard's guinea-pigs, and results like those of MacDougal with plants and of Tower with beetles may lead us to alter the opinion stated. But as it stands now most investigators hold that there are strong general grounds for disbelief in the principle, and also that it lacks experimental proof.
