tively. The segment governing the arms is especially active, in male frogs, in the breeding season; and these members alone with the breast and back appertaining to them, everything else being cut away, will then actively grasp a finger placed between them and remain hanging to it for a considerable time.
The spinal cord in other animals has analogous powers. Even in man it makes movements of defence. Paraplegics draw up their legs when tickled; and Robin, on tickling the breast of a criminal an hour after decapitation, saw the arm and hand move towards the spot. Of the lower functions of the mammalian cord, studied so ably by Goltz and others, this is not the place to speak.
If, in a second animal, the cut be made just behind the optic lobes so that the cerebellum and medulla oblongata remain attached to the cord, then swallowing, breathing, crawling, and a rather enfeebled jumping and swimming are added to the movements previously observed.[1] There are other reflexes too. The animal, thrown on his back, immediately turns over to his belly. Placed in a shallow bowl, which is floated on water and made to rotate, he responds to the rotation by first turning his head and then waltzing around with his entire body, in the opposite direction to the whirling of the bowl. If his support be tilted so that his head points downwards, he points it up; he points it down if it be pointed upwards, to the right if it be pointed to the left, etc. But his reactions do not go farther than these movements of the head. He will not, like frogs whose thalami are preserved, climb up a board if the latter be tilted, but will slide off it to the ground.
If the cut be made on another frog between the thalami and the optic lobes, the locomotion both on land and water becomes quite normal, and, in addition to the reflexes already shown by the lower centres, he croaks regularly whenever he is pinched under the arms. He compensates rotations, etc., by movements of the head, and turns over from his back; but still drops off his tilted
- ↑ It should be said that this particular cut commonly proves fatal. The text refers to the rare cases which survive.
