selves we should call an idea. His reactions to outward stimuli vary their form, too. Instead of making simple defensive movements with his hind legs like a headless frog if touched, or of giving one or two leaps and then sitting still like a hemisphereless one, he makes persistent and varied efforts at escape, as if, not the mere contact of the physiologist's hand, but the notion of danger suggested by it were now his spur. Led by the feeling of hunger, too, he goes in search of insects, fish, or smaller frogs, and varies his procedure with each species of victim. The physiologist cannot by manipulating him elicit croaking, crawling up a board, swimming or stopping, at will. His conduct has become incalculable. We can no longer foretell it exactly. Effort to escape is his dominant reaction, but he may do anything else, even swell up and become perfectly passive in our hands.
Such are the phenomena commonly observed, and such the impressions which one naturally receives. Certain general conclusions follow irresistibly. First of all the following:
The acts of all the centres involve the use of the same muscles. When a headless frog's hind leg wipes the acid, he calls into play all the leg-muscles which a frog with his full medulla oblongata and cerebellum uses when he turns from his back to his belly. Their contractions are, however, combined differently in the two cases, so that the results vary widely. We must consequently conclude that specific arrangements of cells and fibres exist in the cord for wiping, in the medulla for turning over, etc. Similarly they exist in the thalami for jumping over seen obstacles and for balancing the moved body; in the optic lobes for creeping backwards, or what not. But in the hemispheres, since the presence of these organs brings no new elementary form of movement with it, but only determines differently the occasions on which the movements shall occur, making the usual stimuli less fatal and machine-like; we need suppose no such machinery directly co-ordinative of muscular contractions to exist. We may rather assume, when the mandate for a wiping-movement is sent forth by
