admitted that emotion adds to the efficacy of behaviour, and this is the view of Professor Lloyd Morgan. " Whatever may be the exact psychological nature of the emotions, it may be regarded," he says, "as certain that they introduce into the conscious situation elements which contribute not a little to the energy of behaviour. They are important conditions to vigorous and sustained conation." Therefore, if it be true that the female raises the emotional tone of the male, the result will be an increased flow of energy into all the specific modes of behaviour connected with reproduction, amongst which those directly concerned in the securing and defence of the territory will receive their share; so that instead of a progressive weakening of just those elements in the situation which make for success, the level of their efficiency will be maintained as a result of such reinforcement. But the female becomes intolerant of her own sex when she has discovered a male ready to breed, and, later, assists her mate in resisting intrusion; and by raising her emotional tone, he may be the means of furthering more strenuous behaviour on her part. Each member of the pair would in this way contribute towards the energy of behaviour of its mate, and hence add indirectly to the security of the territory.
It may be well to illustrate the foregoing remarks. Suppose that there is a small piece of woodland barely sufficient to hold three pairs of Willow- Warblers, and suppose that the male and female in the middle territory did not respond to
