Page:Territory in Bird Life by Henry Eliot Howard (London, John Murray edition).djvu/258

This page has been proofread, but needs to be validated.
200
TERRITORY AND REPRODUCTION

suitable breeding ground, no greater calamity could befall the species than that some members should exercise dominion over too large an area of the habitable part of the cliff and thus prevent others from breeding. Under such conditions the race could not endure, since in this, as in every case, its survival must depend upon a close correspondence between the behaviour of the individual and the circumstances in the external environment.

Scarcity of suitable cliffs is the principal reason of the ledges being so closely packed with Guillemots, just as it accounts for this part of the precipice being crowded with Kittiwake Gulls, that part with Herring-Gulls, and that part again with Razorbills and Puffins. Yet each individual preserves its few square feet of rock or soil from molestation, and the area each one occupies varies according to the conditions of existence of the species. Thus the Herring-Gull occupies a comparatively small area, although one many times larger than that of the Guillemot. It requires more space than the latter, owing to the fact that it not only builds a nest but rears four instead of a single offspring, and it can be allowed this, because, since its young remain in the nest until they are capable of sustained flight, it can make use of many miles of cliff from which the tide recedes at the base, or which have, at their ba,se, rocks jutting out into the sea; but manifestly it cannot be allowed so much space as the Bunting.