aroused. Thus we come to recognise fear, anger, or sexual emotion, by just the particular sound which is emitted. But even if we are going too far in referring particular calls to particular emotions, we can, without a doubt, divide them into two broad categories—those which are pleasurable and those which are the reverse. And we need have no hesitation in placing the particular call to which I allude in the first of these two categories, not only on account of the nature of the sound produced, but because the activities which are aroused are not such as normally accompany irritation. This is well seen if the behaviour of different individuals be closely observed. After resting on one leg for some time, first one and then another is seized with cramp, and running a few yards in an ungainly way, bumps up against its companions as if it had not full control over its movements. Its behaviour produces irritation which is expressed by a vocal outburst, and followed by actions the meaning of which is clear. Moreover, the call is taken up by other individuals and sweeps over part of the flock as does the greeting. But the nature of the cry is entirely different from that which greets the arrival of a companion—humanly speaking it is a passionate and impatient utterance, the height of displeasure. The arrival, then, acts as a stimulus to something in the inherited constitution which is expressed in, and presumably is satisfied by, this vocal outburst; and, since the bird that arrives joins also in the
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RELATION OF TERRITORY TO MIGRATION
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