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24
Indiana University Studies

is no uniformity in current practice, and the only attempts at uniformity have been based on purely artificial distinctions. The resolution of the terminology must take into account as a question of convenience that no category higher than the genus may be written into the nomenclature, and as a matter of fact that there are often three or four degrees of phylogenetic affinities which may be recognized below what seems to be best called a genus.

The ichthyologists believe that they have the solution in calling this lowest category a species and the second category a genus, thereby making their order the equivalent of many a genus among insects. I interpret the mammalogists to mean that they call a Mendelian race a variety, and the fundamental taxonomic unit a subspecies, which they imply is an incipient “species” (their next category). The botanists, as nearly as I can perceive, call their lowest unit either a variety or a species, depending upon its remoteness from the native heath of the botanist and his field of experience.

I am at a loss for a solution of this difficulty. It seems unreasonable to expect that this first category will ever be called anything but a species by biologists who are not systematists, and in that sense I shall use the word in general discussion in this study. It will be impossible to adopt this meaning in our system of classification without inventing a new name for a category between this and the genus, and I have not the temerity to propose such a name while taxonomists are as far removed from biologic realities as the codes of nomenclature and much current systematics would indicate. Consequently, in the systematic portion of this paper I have adopted the term variety for the category which, after all, fulfills the species concept. I can only plead that I am conscious of the inevitable confusion this involves, and desirous of making amends as soon as some one proposes a solution—but I shall look for a solution that will coördinate biologic concepts of species with questions of convenience in systematic botany and zoölogy.